《云南植物志》|| 茜草科 Rubiaceae ||山花属 Prismatomeris Thw. ||四蕊三角瓣花 (原亚种) Prismatomeris tetrandra (Roxb.) K. Schum. ssp. tetrandra

1a.四蕊三角瓣花（原亚种）
ssp. tetrandra
灌木或小乔木，高可达8米。小枝四棱柱形。叶披针形或长圆状披针形，长8—17厘米，宽3—6厘米，先端渐尖或短尖，基部楔形，两面无毛，侧脉5—9对，弯拱相联，在叶两面凸起；叶柄长5—15毫米；托叶三角形，脱落。花芳香，2—16朵簇生或组成伞形花序，顶生或腋生，总花梗极短；苞片与托叶相似；花梗长5—15毫米；花萼杯状，长2—4毫米，无毛或有疏腺毛，花萼裂片不明显；花冠白色或淡紫色，高脚碟状，长2—4厘米，冠管长约2厘米，花冠裂片4—5，披针形，长约1厘米，镊合状排列；雄蕊4—5，着生在冠管上部，花丝极短或无，花药线形，长3—4毫米，内藏或伸出；花柱常内藏，柱头2裂，长约4毫米，子房2室。核果近球形，直径约8毫米，顶部具环状宿萼；种子1—2颗。 花期5—9月，果期9—12月。
分布于印度和孟加拉。云南不产。

《云南植物志》|| 茜草科 Rubiaceae ||山花属 Prismatomeris Thw. ||多花三角瓣花 (亚种) Prismatomeris tetrandra (Roxb.) K. Schum. ssp. multiflora (Ridl.) Y. Z. Ruan

1b.多花三角瓣花（亚种）（植物分类学报）图版61: 1—3
ssp. multiflora (Ridl.) Y. Z. Ruan (1988); 中国植物志 (1999)*.
Prismatomeris multiflora Ridl. (1940); P. tetrandra auct. non (Roxb.) K. Schum.: 云南种子植物名录 (1984); 云南树木图志 (1991)*.
灌木或小乔木，高可达8米。小枝四棱柱形。叶披针形或长圆状披针形，长8—17厘米，宽3—6厘米，先端渐尖或短尖，基部楔形，两面无毛，侧脉5—9对，弯拱相联，在叶两面凸起；叶柄长5—15毫米；托叶三角形，脱落。花芳香，2—16朵簇生或组成伞形花序，顶生或腋生，总花梗极短；苞片与托叶相似；花梗长5—15毫米；花萼杯状，长2—4毫米，无毛或有疏腺毛，花萼裂片不明显；花冠白色或淡紫色，高脚碟状，长2—4厘米，冠管长约2厘米，花冠裂片4—5，披针形，长约1厘米，镊合状排列；雄蕊4—5，着生在冠管上部，花丝极短或无，花药线形，长3—4毫米，内藏或伸出；花柱常内藏，柱头2裂，长约4毫米，子房2室。核果近球形，直径约8毫米，顶部具环状宿萼；种子1—2颗。 花期5—9月，果期9—12月。
产金平、勐腊、景洪、勐海；生于海拔400—1200米处的山谷林下或灌丛。分布于泰国。

《云南植物志》|| 茜草科 Rubiaceae ||山花属 Prismatomeris Thw. ||四蕊三角瓣花 Prismatomeris tetrandra (Roxb.) K. Schum.

1.四蕊三角瓣花（植物分类学报）
Prismatomeris tetrandra (Roxb.) K. Schum. in Engl. u. Prantl (1891).
灌木或小乔木，高可达8米。小枝四棱柱形。叶披针形或长圆状披针形，长8—17厘米，宽3—6厘米，先端渐尖或短尖，基部楔形，两面无毛，侧脉5—9对，弯拱相联，在叶两面凸起；叶柄长5—15毫米；托叶三角形，脱落。花芳香，2—16朵簇生或组成伞形花序，顶生或腋生，总花梗极短；苞片与托叶相似；花梗长5—15毫米；花萼杯状，长2—4毫米，无毛或有疏腺毛，花萼裂片不明显；花冠白色或淡紫色，高脚碟状，长2—4厘米，冠管长约2厘米，花冠裂片4—5，披针形，长约1厘米，镊合状排列；雄蕊4—5，着生在冠管上部，花丝极短或无，花药线形，长3—4毫米，内藏或伸出；花柱常内藏，柱头2裂，长约4毫米，子房2室。核果近球形，直径约8毫米，顶部具环状宿萼；种子1—2颗。 花期5—9月，果期9—12月。
产金平、勐腊、景洪、勐海；生于海拔400—1200米处的山谷林下或灌丛。分布于泰国。

《Flora of China》|| Rubiaceae || Prismatomeris ||四蕊三角瓣花 Prismatomeris tetrandra

1. Prismatomeris tetrandra (Roxburgh) K. Schumann in Engler & Prantl, Nat. Pflanzenfam. 4(4): 138. 1891.

四蕊三角瓣花 si rui san jiao ban hua

Coffea tetrandra Roxburgh, Fl. Ind. 2: 193. 1824; Octotropis terminalis C. B. Clarke; Prismatomeris connata Y. Z. Ruan; P. connata subsp. hainanensis Y. Z. Ruan; P. multiflora Ridley; P. tetrandra subsp. multiflora (Ridley) Y. Z. Ruan.

Shrubs or small trees, to 8 m tall; branches quadrangular to subterete, glabrous. Petiole 4-15 mm, glabrous; leaf blade drying leathery to stiffly papery and rather shiny, lanceolate, elliptic, elliptic-oblong, ovate, obovate, or oblong-lanceolate, 4-18 × 2-6 cm, glabrous, base cuneate to acute, apex acuminate or acute to obtuse; secondary veins 5-9 pairs; stipules with sheath portion 0.2-1 mm, truncate, persistent or deciduous by fragmentation, bilobed, lobes linear to narrowly triangular, 0.1-0.5 mm, caducous. Peduncles 1-16 and fascicled or umbellate, 5-35 mm, simple or bearing an umbellate group of pedicels, at base with stipuliform bracts; pedicels when present 5-15 mm. Flowers pedunculate or pedicellate. Calyx glabrous or sparsely glandular-puberulent; hypanthium portion hemispherical, 1.5-2 mm; limb 1-2 mm, truncate or 4- or 5-denticulate. Corolla white or pale purple, salverform, glabrous outside; tube 14-20 mm; lobes 4 or 5, lanceolate, 7-10 mm, abaxially (i.e., dorsally) ridged at least in bud, acute to obtuse. Drupes subglobose, 8-12 mm in diam., glabrous, smooth. Fl. May-Sep, fr. Sep-Dec.

Forests, thickets; 300-2400 m. Fujian, Guangdong, Guangxi, Hainan, S Yunnan [Cambodia, India, Thailand, Vietnam].

Johansson (Opera Bot. 94: 29. 1987) recognized two subspecies, Prismatomeris tetrandra subsp. tetrandra and P. tetrandra subsp. malayana (Ridley) J. T. Johannson; he reported only subsp. tetrandra from China. However, Puff et al. (Rubiaceae of Thailand, 118. 2005) noted that further observation shows that even these taxa cannot be completely separated either morphologically or geographically. Y. Z. Ruan (in FRPS 71(2): 178. 1999) included the Chinese plants in P. tetrandra subsp. multiflora, which Johansson synonymized with subsp. tetrandra. Ruan (loc. cit.) reported that the flowers of P. connata are occasionally unisexual; this was not further documented there and has not been reported by others nor confirmed with specimens. Ruan (loc. cit.: 179) also described the calyx lobes as ca. 3 mm, which has not been reported by other authors nor seen on the rather ample materials studied.

This species was newly reported from India by Ayappan and Parthasarathy (J. Econ. Taxon. Bot. 29: 802-804. 2005), and its distribution in Cambodia and Vietnam is here reported based on specimens at P annotated by Johansson after the publication of his article.