《中国植物志》|| 茜草科 RUBIACEAE ||滇丁香属 Luculia Sweet ||馥郁滇丁香 Luculia gratissima (Wall.) R. Sweet

3. 馥郁滇丁香（中国高等植物图鉴）图版60: 10-12
Luculia gratissima (Wall.) Sweet. Brit. Fl. Gard. 2: t. 145. 1826; Hook. Curtis's Bot. Mag. 68: t. 3946. 1842; Brandis, Ind. Trees 373. 1906; Hutchins. in Sargent, Pl. Wils. 3: 407. 1916; Levl. Cat. Pl. Yunnan 246. 1917; How in Sunyatsenia 7 (1-2): 27. 1948, pro min. p.; 中国高等植物图鉴4: 201, 图5816. 1975; Hara et Gould in Hara et al. Enum. Flow. Pl. Nepal 2: 204. 1979; 云南种子植物名录, 下册: 1261. 1984; 西藏植物志4: 429. 1985; H. Koba et al. Name List Flow. Pl. Gymnosp. Nepal 307. 1994. ——Cinchona gratissima Wall. in Roxb. Fl. Ind. ed. Carey 2: 154. 1824.——Mussaenda luculia Buch. -Ham. ex D. Don, Prodr. Fl. Nepal 139. 1825, nom. illeg.
灌木或小乔木，高达5米；树皮薄，淡褐色；小枝有皮孔，幼嫩时淡红色，有疏柔毛。叶纸质或薄革质，卵状长圆形、椭圆形或椭圆状倒披针形，长5-15厘米，宽2-6厘米，顶端渐尖，基部楔形或短尖，全缘，上面无毛，下面沿中脉和侧脉上被柔毛，脉腋内有时有簇毛，其余无毛或有疏柔毛；侧脉8-12对，在下面凸起；叶柄长0.8-2厘米；托叶披针形，早落。伞房状的聚伞花序大，顶生，多花，有多数线形早落的苞片；总花梗被疏柔毛；花大，美丽，芳香；萼管被疏柔毛，裂片披针形，直立，长1-1.6厘米；花冠红色，高脚碟状，长约5厘米，冠管细圆柱形，裂片圆形，开展；雄蕊着生于冠管内，花丝长1.5毫米，花药长5毫米；花柱丝状，柱头大，2裂，内藏，子房倒卵形，有柔毛。蒴果倒卵状长圆形，被疏柔毛或毛以后脱落，长约2厘米。花、果期4-11月。
产于云南西部和西南部，西藏墨脱；生于海拔800-2400米处的山地林中或灌丛中。国外分布于印度东北部，尼泊尔，不丹，缅甸，泰国，越南等地。

《云南植物志》|| 茜草科 Rubiaceae ||滇丁香属 Luculia Sweet ||馥郁滇丁香 Luculia gratissima (Wall.) Sweet

3.馥郁滇丁香（图鉴）
Luculia gratissima (Wall.) Sweet (1826)*; 图鉴 (1975)*; Hara et Gould in Hara et al. (1979); 云南种子植物名录 (1984); 横断山区维管植物 (1994); 中国植物志 (1999)*.
Cinchona gratissima Wall. in Roxb. (1824); Mussaenda luculia Buch.-Ham. ex D. Don (1825), nom. illegit.
灌木或小乔木，高达5米；树皮薄，淡褐色。小枝有皮孔，幼嫩时淡红色，有疏柔毛。叶纸质或薄革质，卵状长圆形、椭圆形或椭圆状倒披针形，长5—15厘米，宽2—6厘米，先端渐尖，基部楔形或短尖，全缘，叶面无毛，背面沿中脉和侧脉上被柔毛，脉腋内有时有簇毛，其余无毛或有疏柔毛，侧脉8—12对，在背面凸起；叶柄长0.8—2厘米；托叶披针形，早落。伞房状的聚伞花序，大，顶生，多花，有多数线形早落的苞片；总花梗被疏柔毛；花大，美丽，芳香；萼管被疏柔毛，萼裂片披针形，直立，长1—1.6厘米；花冠红色，高脚碟状，长5厘米，花冠裂片圆形，开展；雄蕊着生于冠管内，花丝长1.5毫米，花药长5毫米；花柱丝状，柱头大，2裂，内藏，子房倒卵形，有柔毛。蒴果倒卵状长圆形，被疏柔毛或毛以后脱落，长2厘米。花果期4—11月。
产丽江、贡山、福贡、泸水、大理、潞西、思茅、景洪、腾冲；生于海拔1350—3500米处的山谷溪边林中或灌丛中。西藏也有。分布于越南、泰国、缅甸、不丹、锡金、尼泊尔、孟加拉国、印度。

《Flora of China》|| Rubiaceae || Luculia ||馥郁滇丁香 Luculia gratissima

1. Luculia gratissima (Wallich) Sweet, Brit. Fl. Gard. 2: t. 145. 1826.

馥郁滇丁香 fu yu dian ding xiang

Cinchona gratissima Wallich in Roxburgh, Fl. Ind. 2: 154. 1824; Mussaenda luculia Buchanan-Hamilton ex D. Don, nom. illeg. superfl.

Shrubs or small trees, to 5 m tall, with thin pale brown bark; branches somewhat flattened to subterete, pilosulous [to glabrous], sometimes with sparse elliptic lenticels. Petiole 0.8-2 cm, pilosulous to glabrous; leaf blade drying papery or thinly leathery, elliptic, lanceolate-elliptic, lanceolate, or elliptic-oblong, 5-15 × 2-6 cm, adaxially glabrous, abaxially pilosulous at least along principal veins, base cuneate or acute, apex acute to acuminate; secondary veins 8-12 pairs, sometimes with pilosulous domatia; stipules lanceolate to narrowly lanceolate, 6-8 mm, glabrous to strigillose, acute to acuminate. Inflorescence corymbiform, many flowered, villosulous to pilosulous; peduncle 2-3 cm; branched portion 3-5 × 6-8 cm (not including corollas); bracts linear to narrowly ligulate, 4-7 mm, acute; pedicels 4-5 mm. Calyx glabrous to densely hirtellous; ovary portion obovoid, 3-4 mm; lobes lanceolate, oblanceolate, or narrowly ligulate, 10-16 mm, acute. Corolla red, outside glabrous; tube 30-50 mm, slenderly cylindrical; lobes suborbicular to broadly elliptic, 11-12 × 12-15 mm, rounded. Capsule 1-2 × 1-1.5 cm, pilosulous to glabrous; seeds 2-3 mm, spongy-reticulate. Fl. and fr. Apr-Nov.

Forests or thickets on mountains; 800-2400 m. Xizang (Mêdog), Yunnan [Bhutan, NE India, Myanmar, Nepal, Thailand, Vietnam].

W. C. Chen (in FRPS 71(1): 239. 1999) distinguished this from other species of Luculia in part by its villous pubescence on the calyx and hypanthium, but some individual specimens seen vary from densely pubescent to glabrous. The pollination biology of this species was studied by Murray (Ann. Bot. 65: 691-698. 1990).