《中国植物志》|| 菊科 COMPOSITAE ||珠光系 Ser. Margaripes (DC.) Boiss. ||珠光香青 Anaphalis margaritacea (L.) Benth. & Hook.f.

3. 珠光香青 山萩
Anaphalis margaritacea (L.) Benth. et Hook. f., Gen. Pl. 2: 303. 1862. ——Gnaphalium margaritaceum L. Sp. Pl. 850. 1753.
根状茎横走或斜升，木质，有具褐色鳞片的短匍枝。茎直立或斜升，单生或少数丛生，高30-60稀达100厘米，常粗壮，不分枝，稀在断茎或健株上有分枝，被灰白色棉毛，下部木质。下部叶在花期常枯萎，顶端钝；中部叶开展，线形或线状披针形，长5-9厘米，宽0.3-1.2厘米，稀更宽，基部稍狭或急狭，多少抱茎，不下延，边缘平，顶端渐尖，有小尖头，上部叶渐小，有长尖头，全部叶稍革质，上面被蛛丝状毛，下面被灰白色至红褐色厚棉毛，有单脉或3-5出脉。头状花序多数，在茎和枝端排列成复伞房状，稀较少而排列成伞房状；花序梗长4-17毫米。总苞宽钟状或半球状，长5-8毫米，径8-13毫米；总苞片5-7层，多少开展，基部多少褐色，上部白色，外层长达总苞全长的三分之一，卵圆形，被棉毛，内层卵圆至长椭圆形，长5毫米，宽2.5毫米，在雄株宽达3毫米，顶端圆形或稍尖，最内层线状倒披针形，宽0.5毫米，有长达全长四分之三的爪部。花托蜂窝状。雌株头状花序外围有多层雌花，中央有3-20雄花；雄株头状花全部有雄花或外围有极少数雌花。花冠长3-5毫米。冠毛较花冠稍长，在雌花细丝状；在雄花上部较粗厚，有细锯齿。瘦果长椭圆形，长0.7毫米，有小腺点。花果期8-11月。
广泛分布于我国西南部、西部、中部和印度、日本、朝鲜、苏联远东地区、美洲北部，在欧洲驯化或栽培。

《云南植物志》|| 菊科 Compositae ||珠光香青 Anaphalis margaritacea (L.) Benth. ||珠光香青 (原变种) Anaphalis margaritacea (L.) Benth. var. margaritacea

la. 珠光香青 (原变种)
Anaphalis margaritacea (L.) Benth. var. margaritacea
多年生草本，高30-90厘米。根茎粗或细，垂直，稀斜生或横走，有短匍匐枝；主根长达8厘米，粗达4毫米，具侧根和多数纤维状细根。茎直立或基部斜生，单生或少数簇生，基部粗2-5毫米，不分枝，被灰白色绵毛或蛛丝状毛。茎下部叶在开花期枯萎，中部叶线状披针形，长4-9厘米，宽4-8毫米，先端渐尖，具尖头，基部渐狭，不下延，边缘全缘，平或稍反卷，表面灰绿色，被蛛丝状毛，后常脱毛，背面被灰白色、淡褐色或红褐色厚绵毛，中脉在背面凸起，有时具1对细侧脉，无叶柄，上部叶渐小。头状花序多数，于茎顶排列成复伞房状花序；花序梗长3-15毫米，被灰白色绵毛；总苞宽钟形或半球形，长6-8毫米，径8-13毫米；总苞片5-7层，略开展，外层卵圆形，长2-3毫米，背面被灰白色蛛丝状毛，内层长卵形或长椭圆形，长6-8毫米，全部总苞片白色，基部浅褐色，干膜质，先端钝或急尖，常具爪。雌株的头状花序外围有多层雌花，中央有3-20朵雄花；雄株的头状花序全为雄花或外围有少数雌花。雌花花冠毛管状，长3-5毫米；雄花花冠管状，长3-5毫米，管部稍长于檐部。瘦果狭长圆形，长0.5-0.7毫米，具细腺点；冠毛稍长于花冠，雌花的冠毛丝状，雄花的冠毛上部稍增粗。花果期7-11月。
我省大部分地区有分布；生于海拔 (900-) 1200-3000 (-3600) 米的林下、林缘、灌丛中、山坡草地或荒地和路边。江西、台湾、河南、湖北、湖南、广西、陕西、甘肃、青海、四川、贵州和西藏有分布。不丹、锡金、尼泊尔、印度北部、中南半岛、俄罗斯、日本及北美也有。
全草药用，清热泻火、燥湿。茎叶和花含芳香油，可作调香原料。

《云南植物志》|| 菊科 Compositae ||香青属 Anaphalis DC. ||珠光香青 Anaphalis margaritacea (L.) Benth.

1. 珠光香青 (图鉴)
火草 (维西、腾冲、河口、东川) ，火油草 (河口) ，小火草、大火草 (贵州) ，山萩
Anaphalis margaritacea (L.) Benth. in Benth. et Hook. f. (1862), (1873); C. B. Clarke (1876); Maxim. (1881); Diels (1900), (1907); Komarov (1907); Rehd. et Kobuski (1933); Hand.-Mazz. (1936), (1938); Kitam. (1937), (1961), (1968), in Hara (1971), (1975); S. Y. Hu (1965); Chen, Ling et al. (1966); 图鉴 (1975)*; 中国植物志 (1979); A. O. Chater in Hara et al. (1982); 云南种子植物名录 (1984); 西藏植物志 (1985); 秦岭植物志 (1985)*; 贵州植物志 (1989); 横断山区维管植物 (1994).
Gnaphalium margaritaceum L. (1753); Helichrysum margaritaceum Moench. (1794); Antennaria margaritacea R. Br. (1817), nom., et in DC. (1837); Maxim. (1859); A. cinnamomea DC. var. angustior Miq. (1866); Anaphalis cinnamomea DC. var. angustior Nakai (1926); A. margaritacea (L.) Benth. ssp. angustior (Miq.) Kitam. (1937); S. Y. Hu (1965); A. griffithii auct. non Hook. f.: Diels (1912).
多年生草本，高30-90厘米。根茎粗或细，垂直，稀斜生或横走，有短匍匐枝；主根长达8厘米，粗达4毫米，具侧根和多数纤维状细根。茎直立或基部斜生，单生或少数簇生，基部粗2-5毫米，不分枝，被灰白色绵毛或蛛丝状毛。茎下部叶在开花期枯萎，中部叶线状披针形，长4-9厘米，宽4-8毫米，先端渐尖，具尖头，基部渐狭，不下延，边缘全缘，平或稍反卷，表面灰绿色，被蛛丝状毛，后常脱毛，背面被灰白色、淡褐色或红褐色厚绵毛，中脉在背面凸起，有时具1对细侧脉，无叶柄，上部叶渐小。头状花序多数，于茎顶排列成复伞房状花序；花序梗长3-15毫米，被灰白色绵毛；总苞宽钟形或半球形，长6-8毫米，径8-13毫米；总苞片5-7层，略开展，外层卵圆形，长2-3毫米，背面被灰白色蛛丝状毛，内层长卵形或长椭圆形，长6-8毫米，全部总苞片白色，基部浅褐色，干膜质，先端钝或急尖，常具爪。雌株的头状花序外围有多层雌花，中央有3-20朵雄花；雄株的头状花序全为雄花或外围有少数雌花。雌花花冠毛管状，长3-5毫米；雄花花冠管状，长3-5毫米，管部稍长于檐部。瘦果狭长圆形，长0.5-0.7毫米，具细腺点；冠毛稍长于花冠，雌花的冠毛丝状，雄花的冠毛上部稍增粗。花果期7-11月。
我省大部分地区有分布；生于海拔 (900-) 1200-3000 (-3600) 米的林下、林缘、灌丛中、山坡草地或荒地和路边。江西、台湾、河南、湖北、湖南、广西、陕西、甘肃、青海、四川、贵州和西藏有分布。不丹、锡金、尼泊尔、印度北部、中南半岛、俄罗斯、日本及北美也有。
全草药用，清热泻火、燥湿。茎叶和花含芳香油，可作调香原料。

《Flora of North America》|| Asteraceae || Anaphalis || Anaphalis margaritacea

1. Anaphalis margaritacea (Linnaeus) Bentham & Hooker f., Gen. Pl. 2: 303. 1873.

Pearly everlasting, anaphale marguerite, immortelle blanche

Gnaphalium margaritaceum Linnaeus, Sp. Pl. 2: 850. 1753; Anaphalis margaritacea var. occidentalis Greene; A. margaritacea var. subalpina (A. Gray) A. Gray

Perennials; rhizomes relatively slender. Stems white, densely and closely tomentose, not glandular. Leaf blades 1–3-nerved, 3–10(–15) cm, bases subclasping, decurrent, margins revolute, abaxial faces tomentose or glabrescent (proximal leaves), not glandular or very sparsely and inconspicuously glandular, adaxial faces green, glabrate. Involucres 5–7 × 6–8(–10) mm. Phyllaries ovate to nearly linear (innermost), subequal to unequal, apices white, opaque. Cypselae 0.5–1 mm, bases constricted into stipiform carpopodia. 2n = 28.

Flowering Jul–Oct (sporadically longer). Dry woods, often with aspen or mixed conifer-hardwood, borders and trails, dunes, fields, roadsides, other open, often disturbed sites; 0–3200 m; St. Pierre and Miquelon; Alta., B.C. , Man., N.B., Nfld. and Labr., N.W.T., N.S., Ont., P.E.I. Que., Sask., Yukon; Alaska, Ariz., Ark., Calif., Colo., Conn., Del., Idaho, Ill., Ind., Iowa, Ky., Maine, Md., Mass., Mich., Minn., Mont., Nebr., Nev., N.H., N.J., N.Mex., N.Y., N.C., Ohio, Oreg., Pa., R.I., S.Dak., Tenn., Tex., Utah, Vt., Va., Wash., W.Va., Wis., Wyo.; Mexico (Baja California); Asia; introduced in Europe.

Anaphalis margaretacea was widely planted as an ornamental and escaped. It apparently naturalized from its native range in both Asia and North America; it is cultivated and naturalized in Europe.

Anaphalis margaritacea has the aspect of Pseudognaphalium; it differs in being subdioecious (polygamo-dioecious; the heads either staminate or primarily pistillate) and in its distinctive cypselar vestiture. It is further recognized by its combination of rhizomatous habit, subclasping-decurrent, bicolor, revolute leaves, and distally white phyllaries. Segregate species and varieties have been described among the North American plants (in addition to the two cited above), based on variation in habit, vestiture, and leaf morphology and density, but the variants appear to be more like a complex series of ecotypes rather than broader evolutionary entities.

《Flora of China》|| Asteraceae || Anaphalis ||珠光香青 Anaphalis margaritacea

3. Anaphalis margaritacea (Linnaeus) Bentham & J. D. Hooker, Gen. Pl. 2: 303. 1873.

珠光香青 zhu guang xiang qing

Herbs, perennial. Rhizome repent, woody, with short brown squamate runners. Stems solitary or a few caespitose, erect or ascending, 30-60(-100) cm, usually thickish, unbranched, rarely branching on broken or normal stems, ash-gray cottony tomentose, base woody. Lower leaves withered by anthesis, apex obtuse; middle leaves expanding, linear or linear-lanceolate, 5-10 cm × 3-12 mm, rarely wider, base attenuate or sharply narrowed, somewhat amplexicaul, margin flat, apex acuminate with small tip; upper leaves gradually smaller with long tip; all leaves slightly leathery, abaxially densely ash-gray to reddish brown lanuginous, adaxially arachnoid or later glabrous, 1-veined, or 3- or 5-veined. Capitula numerous, arranged in compound corymbiform (or corymbiform) synflorescences; peduncle 4-17 mm. Involucre broadly campanulate or semispherical, 5-8 × 8-13 mm; phyllaries 5-7-seriate, somewhat expanding, upper parts white; outer phyllaries up to 1/3 of involucre, ovate, cottony tomentose; middle ones ovate to oblong, ca. 5 × 2.5 mm, up to 3 mm wide in male plants, apex rounded or acuminate; innermost ones linear-lanceolate, ca. 0.5 mm wide, with a claw up to 3/4 of full length. Receptacle alveolate. Predominantly female capitula with many florets, with 3-20 central male florets and many rows of marginal female florets. Predominantly male capitula with many florets, with all central male florets or few rows of marginal female florets. Corolla 3-5 mm. Pappus slightly exserted from corolla, filiform in female florets, incrassate at tip of apex, serrulate in male florets. Achene oblong, ca. 0.7 mm, with glandular dots. Fl. and fr. Jul-Nov. 2n = 28*, 42, 56*.

Conifer and Betula forests, dry soils, subalpine or low mountain grasslands or shrublands, rocky valleys and roadsides, slopes, banks; 300-3400 m. E, S, and W Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, E and W Hubei, Hunan, Jiangxi, Qinghai, S Shanxi, Sichuan, Taiwan, Xizang, Yunnan [Bhutan, India, Japan, Korea, Myanmar, Nepal, Russia (Far East), N Thailand, Vietnam; North America; widely introduced in Europe].