《中国植物志》|| 忍冬科 CAPRIFOLIACEAE ||大叶组 Sect. 7. Megalotinus (Maxim.) Rehd. ||锥序荚蒾 Viburnum pyramidatum Rehder

49. 锥序荚蒾（中国高等植物图鉴）尖锥荚蒾（拉汉种子植物名称）图版14：5-8
Viburnum pyramidatum Rehd. in Sarg. Trees and Shrubs 2: 93. 1908; 中国高等植物图鉴4: 315. 1975.
灌木或小乔木，高达7米，幼枝、芽、叶下面、叶柄和花序均被由黄褐色簇状毛组成的绒毛；当年小枝圆筒状，散生皮孔，二年生小枝变灰黄色，近无毛。叶厚纸质，卵状矩圆形至矩圆形或宽椭圆形，长8-16 (-20) 厘米，顶端渐尖，基部狭窄或近圆形，边缘有牙齿状锯齿或小锯齿，上面有光泽，除中脉初时散生簇状毛外均无毛，侧脉6-7对，弧形，连同中脉下面凸起，小脉横列，下面不明显或略显著；叶柄长1.5-3厘米。圆锥式花序尖塔形，长5-10厘米，直径几相等，由2-4层伞形花序组成，每层具4-6条辐射枝，总花梗长2-4厘米，花生于序轴的第三级辐射枝上；萼筒倒圆锥形，无毛，萼齿三角形，长约为筒的1/3，疏具缘毛；花冠白色，辐状，直径约4毫米，略被簇状短毛，裂片卵形，顶钝圆，长约为筒的两倍；雄蕊略短于花冠，花药宽椭圆形；花柱高出萼齿。果实深红色，矩圆形、宽椭圆形至倒卵状矩圆形，长7-10毫米，直径4-5毫米；核稍扁，具2条深背沟和1条浅腹沟。花期12-1月，果熟期11-12月。
产广西西部和云南东南部。生于山谷疏林或灌丛中，亦见于竹林中或荒芜地，海拔120-1400米。越南北部也有分布。模式标本采自云南蒙自。
本种同侧花组 Sect. Platyphylla Hsu 的广叶荚蒾 V. amplifolium Rehd. 和侧花荚蒾 V. laterale Rehd. 都有关联，但后两种叶的侧脉至少部分伸至齿端，聚伞花序伞房式，生于具1对叶的短枝之顶，与本种区别明显。

《云南植物志》|| 忍冬科 Caprifoliaceae nom. conserv. ||荚蒾属 Viburnum L. ||锥序荚蒾 Viburnum pyramidatum Rehd. in Sarg.

31.锥序荚蒾（图鉴）
黑皮树（马关），铜虾果（河口）
Viburnum pyramidatum Rehd. in Sarg. (1908), (1911); 图鉴 (1975), in nota.
常绿灌木至乔木，高2—7米；当年生枝黄褐色，被簇状毛，散生皮孔，一年生枝变无毛，灰褐色，密生皮孔；冬芽有2鳞片，鳞片密被黄褐色簇状毛。叶片坚纸质，卵状长圆形或长圆形，长8—19厘米，宽4—8厘米，先端渐尖，基部楔形，边缘在基部以上有小牙齿状锯齿，叶面暗绿色，光亮，无毛，背面淡绿色，全面尤其是沿脉上密被簇状微柔毛，侧脉每边6—7条，拱弯，近叶缘网结，与中脉在叶面凹陷，在背面突起，细脉横向平行，背面明显；叶柄长1.5—2.5厘米，腹面具槽，密被簇状微柔毛。圆锥花序尖塔形，由数层伞形花序组成，长5—8厘米，密被黄褐色簇状柔毛，总梗长4—6厘米，伞形花序具4—6条第一级辐射枝；苞片和小苞片长卵形，先端钝，外被黄褐色簇状柔毛；花甚小，着生于伞形花序的第三级辐射枝上；萼筒圆柱形，长1.5毫米，无毛，萼齿卵状圆形，长0.7—1毫米，疏被纤毛；花冠白色，辐状，长约2毫米，直径4毫米，通常无毛，花冠裂片卵状圆形，略长于冠筒。核果长圆形，长约1厘米，宽4—5毫米，红色；核扁而略弯，背具1、腹具2浅槽。花期11—12月，果期3—10月。
产蒙自、金平、河口、西畴、麻栗坡、马关，生于海拔120—1400米的山坡路旁、疏林内或灌丛中；分布于广西西南部。越南北部也有。模式标本采自蒙自。

《Flora of China》|| Adoxaceae || Viburnum ||锥序荚蒾 Viburnum pyramidatum

48. Viburnum pyramidatum Rehder in Sargent, Trees & Shrubs. 2: 93. 1908.

锥序荚蒾 zhui xu jia mi

Shrubs or small shrubs, evergreen, to 7 m tall. Bark gray-brownish. Branchlets of current year brownish, sparsely lenticellate; branchlets of previous year gray-yellowish, terete, subglabrous, with sparse, small, rounded lenticels. Winter buds ovoid-lanceolate, with a pair of separate scales; scales yellow-brown stellate-tomentose. Leaves always opposite, not clustered at apices of branchlets; stipules absent; petiole green, slender, 1.5-3 cm, yellow-brown stellate-tomentose; leaf blade yellowish green when young, ovate-oblong to oblong or broadly elliptic, 8-16(-20) × 4-8 cm, thickly papery, abaxially yellow-brown stellate-tomentose, adaxially lustrous, glabrous throughout except on midvein sparsely stellate-pubescent, midvein raised abaxially, lateral veins 6- or 7-jugate, pinnate, arched, rarely branched, anastomosing near margin, raised abaxially, slightly impressed adaxially, veinlets transverse, inconspicuous or slightly conspicuous abaxially, inconspicuous adaxially, not lobed, base narrowed or subrounded, without glands, margin serrate, apex acuminate. Flowers appearing after leaves; inflorescence a pyramidal panicle, terminal, 5-10 × 5-10 cm; rays whorled; first node of inflorescence with 4-6 rays, dense, densely yellowish brown stellate-pubescent, without large sterile radiant flowers; peduncles 2-4 cm; bracts caducous, leaflike, green, linear-lanceolate, stellate pubescent; bracteoles scalelike. Flowers on rays of 3rd order, not fragrant, sessile or shortly pedicellate. Calyx yellowish green; tube obconical, ca. 1.5 mm, glabrous; lobes triangular, ca. 1/3 as long as calyx tube, sparsely ciliate, apex obtuse. Corolla white, rotate, ca. 4 mm in diam., sparsely stellate-pubescent; tube ca. 1 mm; lobes spreading, ovate, ca. 2 × as long as tube, apex rounded, margin entire. Stamens slightly shorter than corolla, inserted at base of corolla; filaments ca. 2 mm; anthers yellow, broadly elliptic, ca. 1 mm. Styles exceeding calyx lobes; stigmas capitate. Fruit maturing intense red, oblong or broadly ellipsoid to obovoid-oblong, 7-10 × 4-5 mm, base rounded, apex acute, glabrous; pyrenes slightly compressed, oblong, ca. 7 × 4 mm, with 2 deep dorsal grooves and 1 shallow ventral groove, apex rounded. Fl. Nov-Dec, fr. Mar-Oct.

Sparse forests, scrub; 100-1400 m. SW Guangxi, SE Yunnan [N Vietnam].