《中国植物志》|| 岩蕨科 WOODSIACEAE ||岩蕨科 WOODSIACEAE ||膀胱蕨属 Protowoodsia Ching

1. 膀胱蕨属 Protowoodsia Ching
Ching in Sunyatsenia 5 (4): 245. 1940 et in Lingnan Sci. Journ. 21(1-4): 36. 1945 et in Acta Phytotax. Sinica 16 (3): 14. 1978; S. H. Wu et Ching, Fern Fam. ＆ Gen. of China 397, f. 5-143. 1991. ——Physematium Kaulf. Flora 341. 1829; Nakai in Bot. Mag. Tokyo 39: 176. 1925. ——Woodsia subgen. Physematium ＆ subgen. Perrinia Hook. Sp. Fil. 1: 60, 62. 1846. ——Woodsia sect. Physematium Hook. et Bak. Syn. Fil. 46. 1867; Diels in Engl. u. Prantl, Nat, Pflanzenfam. 1 (4): 161. 1899. ——Woodsia Cop. Gen. Fil. 105. 1947, pro parte.
石生小型草本。根状茎直立，先端密被鳞片；鳞片卵状披针形或披针形，棕色，有光泽，全缘。叶簇生或近簇生，通常披散；柄短，无关节，质脆易断而下部宿存；叶片披针形、远长于叶柄，向基部变狭，二回羽状深裂，羽片多数，无毛或略被腺毛，不具鳞片。叶脉分离，羽状，小脉不达叶边。叶质薄，草质，叶轴质脆易断，略被短腺毛。孢子囊群小，圆形，由6-12个孢子囊组成，位于小脉的顶部或中部；囊托隆起，明显；囊群盖大，下位，球圆形或膀胱形，膜质，完全包被孢子囊群，成熟时从顶部开裂，裂口边缘不具睫毛，宿存，或以后裂为瓣状而脱落；孢子囊大，球形，环带纵向，由16-22个增厚细胞组成，孢子囊柄短而细。孢子椭圆形，周壁具褶皱，外壁表面光滑。染色体x=11, (33)。
约12种，分布于北半球温带，有几种分布至南美洲，非洲有1种。我国有下列1种，产于海拔较高的山地。
本属长期未为人们接受，一般的处理方法都是将这群植物作为广义的岩蕨属内的亚组或组。其实在叶柄无关节、球圆形或膀胱形的囊群盖等方面就与岩蕨属截然不同。1925年Nakai详述了它与岩蕨属的差异，膀胱蕨属才逐渐为人注意。
本志采用的膀胱蕨属的含义比Kaulfuss的较为广泛，笔者将Hooker在Woodsia属内建立的Subgen. Perrinia 改隶本属内成为另一组，因为从囊群盖的特征来看，Perrinia与Physematium是一致的。

《Flora of China》|| Woodsiaceae || ||膀胱蕨属 Protowoodsia

1. Protowoodsia Ching, Lingnan Sci. J. 21: 36. 1945.

膀胱蕨属 pang guang jue shu

Plants epilithic, small. Rhizomes erect, short, apex densely scaly; scales brown, lustrous, ovate-lanceolate or lanceolate, entire. Fronds clustered or subclustered, usually diverging; stipe short, not articulate, brittle, usually lower parts persistent; lamina 1-pinnate-pinnatifid, lanceolate, much longer than stipe, herbaceous, glabrous or glandular hairs sparse. Veins free, pinnate, not reaching laminar margin. Sori small, orbicular, consisting of 6-12 sporangia, medial or submarginal; indusia inferior, spherical or bladderlike, completely wrapping sporangia, tearing apically at maturity. Spores ellipsoid, monolete, perispores corrugate, exospore glabrous on surface. x = 33.

One species: Asia.

《Flora of China》|| Woodsiaceae || ||膀胱蕨属 Protowoodsia

1. Protowoodsia Ching, Lingnan Sci. J. 21: 36. 1945.

膀胱蕨属 pang guang jue shu

Plants epilithic, small. Rhizomes erect, short, apex densely scaly; scales brown, lustrous, ovate-lanceolate or lanceolate, entire. Fronds clustered or subclustered, usually diverging; stipe short, not articulate, brittle, usually lower parts persistent; lamina 1-pinnate-pinnatifid, lanceolate, much longer than stipe, herbaceous, glabrous or glandular hairs sparse. Veins free, pinnate, not reaching laminar margin. Sori small, orbicular, consisting of 6-12 sporangia, medial or submarginal; indusia inferior, spherical or bladderlike, completely wrapping sporangia, tearing apically at maturity. Spores ellipsoid, monolete, perispores corrugate, exospore glabrous on surface. x = 33.

One species: Asia.