《中国植物志》|| 水龙骨科 POLYPODIACEAE ||瓦韦组 Sect. Lepisorus ||棕鳞瓦韦 Lepisorus scolopendrium (Buch.-Ham. ex D. Don) Mehra & Bir

35.棕鳞瓦韦 (西藏植物志) 图版14: 5-8
Lepisorus scolopendrium (Ham. ex. D. Don) Menhra et Bir, Res. Bull. Panjab. Univ. Sci. n. s. 15: 168. 1965; F. M. Jarrett, Ind. Fil. Suppl. 101. 1961-1975; Tagawa et K. Iwats. in Fl. Tailand 3 (4): 511. f. 51: 6. 1989; Ching et al. in C. Y. Wu, Fl. Xizang. 1: 304-305. f. 77: 6-10. 1982; Shing in W. T. Wang, Vasc. Hengduan Mts. 1: 165. 1993; S. L. Yu et al. in Bull. Bot. Res. 16 (1): 24. 1996. ——Polypodium scolopendrium Ham. apud. D. Don. Prod., Fl. Nep. 1. 1825. ——Lepisorus excavatus var. scolopendrium (Ham.) Ching in Bull. Fan Mem. In-st. Biol. 4: 69. 1933.
植株高15-30厘米。根状茎横走，粗壮，密被鳞片；鳞片披针形，棕色，网眼近方形，透明，渐尖头，全缘。叶远生或近生；叶柄长2-5厘米，基部疏被鳞片，禾秆色；叶片狭长披针形，长15-45厘米，下部近1/3处为最宽，约1-4厘米，急尖头或尾状渐尖头，边缘近平直或微波状，干后两面呈淡红棕色，草质或薄纸质。主脉上下均隆起，小脉略可见。孢子囊群圆形或椭圆形，通常聚生于叶片上半部，位于主脉和叶边之间，较靠近主脉，彼此相距约等于1-2个孢子囊群体积，幼时被隔丝覆盖；隔丝淡棕色，圆形，全缘。
产台湾（台北）、海南（五指山）、四川（米易）、贵州（凯里）、云南（漾濞、景洪、双柏、洱源、文山、大理、马关、大姚、盈江、屏边、蒙自、贡山、福贡、景东宾川）、西藏（车文、麦通、错那、定结、波密、吉隆、聂拉木的樟木）。附生林下树于或岩石上，海拔500-2800米。尼泊尔和印度北部也产。

《Flora of China》|| Polypodiaceae || Lepisorus ||棕鳞瓦韦 Lepisorus scolopendrium

37. Lepisorus scolopendrium (Buchanan-Hamilton ex D.Don) Mehra & Bir, Res. Bull. Panjab Univ. Sci., n.s. 15: 168. 1964.

棕鳞瓦韦 zong lin wa wei

Polypodium scolopendrium Buchanan-Hamilton ex D. Don, Prodr. Fl. Nepal. 1. 1825; Lepisorus excavatus (Bory ex Willdenow) Ching var. scolopendrium (Buchanan-Hamilton ex D.Don) Ching; Drynaria sesquipedalis J. Smith, nom. illeg. superfl.; Lepisorus paleparaphysus Y. X. Lin; L. sesquipedalis Fraser-Jenkins, nom. illeg. superfl.; L. virescens Ching & S. K. Wu; Pleopeltis scolopendrium (Buchanan-Hamilton ex D.Don) Alston & Bonner; Polypodium excavatum Bory ex Willdenow var.concolor Takeda.

Plants 15-30 cm tall. Rhizomes creeping, strong, 3-6 mm in diam., densely scaly; scales appressed, brown, lanceolate, 4-6 × 0.8-3 mm, thin in texture, margin entire, apex acuminate; lumina subsquare and transparent. Fronds 0.2-2 cm apart; stipe straw-colored, 2-3(-5) cm, 1-2 mm in diam., sparsely scaly at base; lamina reddish brown on both surfaces when dried, narrowly lanceolate, 12-45 × (1-)1.5-4.5 cm, widest at or below middle, herbaceous or thinly papery, abaxially with small, lanceolate, scales on costa, bicolored, margin ± flat, straight or slightly undulate, apex acute or long caudate-acuminate; costa raised on both sides, veinlets slightly visible. Sori usually along distal half of lamina, close to costa, orbicular or elliptic, 2-3 mm in diam.; paraphyses usually brown, sometimes colorless on immature fronds, orbicular, 0.5-0.7 mm in diam., margin entire.

On tree trunks or rocks in forests; (500-)1400-2800(-3200) m. Guizhou, Hainan, Sichuan, ?Taiwan, Xizang, Yunnan [Bhutan, N India, Nepal].

The original name, Polypodium scolopendrium, must be considered a homonym of P. scolopendria, the basionym of Phymatosorus scolopendria (N. L. Burman) Pichi Sermolli. The epithet was validated at varietal rank by Ching and later elevated to species rank by Mehra and Bir. Smith published Drynaria sesquipedalis as an intended new name for Polypodium scolopendrium Buchanan-Hamilton ex D. Don, not of N. L. Burman, but included Pleopeltis nuda Hooker, thus making it illegitimate. Fraser-Jenkins missed this when publishing Lepisorus sesquipedalis to use for this taxon. Bir and Trikha (Amer. Fern J. 64(2): 54. 1974) chose L. leiopteris (Kunze) Bir & Trikha as the name for this taxon, but Fraser-Jenkins (Taxon. Revis. Indian Subcontinental Pteridophytes, 38. 2008) claims that Polypodium leiopteris is a synonym of L. nudus.

Lepisorus virescens was separated from L. scolopendrium by having the lamina less than 20 cm, thin in texture, and with narrowly attenuate base merging into an indistinct stipe and the plagiotropic, oblong sori, closer to the costa than to the margin. We checked more specimens and observed living plants and found that there is considerable variation in these leaf characters, such that they cannot be used to distinguish taxa. Lepisorus virescens is just a juvenile form of L. scolopendrium. Lepisorus paleparaphysus was distinguished from L. scolopendrium by the nearly colorless paraphyses. We checked more specimens and found colorless paraphyses present within other species of this section, especially in immature laminae.