《中国植物志》|| 小檗科 BERBERIDACEAE ||小檗属 Berberis Linn. ||有棱小檗 Berberis angulosa Wall. ex Hook.f. & Thoms.

8．有棱小檗
Berberis angulosa Wall. ex Hook. f. et Thoms. Fl. Ind. 1: 227, 1855; et in Bot. Mag. 115: t. 7071, 1889; Ahrendt in Journ. Linn. Soc. Bot. 57: 113, 1961; 西藏植物志 2: 146, 图 44, 1-7, 1985.
落叶灌木，高0.5-2米。老枝被柔毛，暗褐色，具深槽，幼枝黄棕色，被微柔毛；茎刺细弱，单生或三分叉，有时五分叉，长7-12毫米，有时被微毛。叶纸质，倒卵形，长1.5-2.5厘米，宽7-14毫米，先端钝形或急尖，基部楔形，上面亮深黄绿色，背面淡黄绿色有光泽，两面叶脉不显，无毛，叶缘平展，有时被微毛，全缘。花单生；黄色；花梗长3-5毫米，下垂，被短柔毛；小苞片卵形、先端渐尖，长约3.5毫米，宽约2毫米；萼片2轮，外萼片椭圆形，长约5.5毫米，宽约3.5毫米，内萼片倒卵形，长约9.2毫米，宽约6.1毫米；花瓣倒卵形，长约6毫米，宽约4.8毫米，先端近全缘，基部溢缩呈爪，具2枚分离的长圆形腺体；雄蕊长约4毫米，药隔稍延伸，先端圆钝；子房含胚珠6枚，胚珠近无柄。浆果近球形，亮红色，长10-12毫米，直径9-12毫米，顶端具不明显宿存花柱或无宿存花柱，不被白粉。花期5-6月，果期7-8月。
产于西藏、青海。生于疏林、灌丛中或灌丛草地。海拔3500-4500米。锡金、尼泊尔和印度东北部也有分布。模式标本采自尼泊尔。在我国西藏聂拉木地区为高山灌丛优势种。

《Flora of China》|| Berberidaceae || Berberis ||有棱小檗 Berberis angulosa

8. Berberis angulosa Wallich ex J. D. Hooker & Thomson, Fl. Ind. 1: 227. 1855.

有棱小檗 you leng xiao bo

Shrubs, deciduous, to 2 m tall. Branches dark brown, pubescent, sulcate; shoots yellow-brown, puberulous; spines simple or 3(or 5)-fid, 7-12 mm, slender, rarely puberulous. Leaf blade abaxially shiny, pale yellow-green, adaxially shiny, yellow-green, obovate, 1.5-2.5 × 0.7-1.4 cm, papery, both surfaces glabrous, inconspicuously veined, base cuneate, margin entire, apex obtuse or acute. Flowers solitary. Pedicels pendulous, 3-5 mm, pubescent; bracteoles ovate, ca. 3.5 × 2 mm, apex acuminate. Sepals in 2 whorls; outer sepals elliptic, ca. 5.5 × 3.5 mm; inner sepals obovate, ca. 9.2 × 6.1 mm. Petals obovate, ca. 6 × 4.8 mm, base clawed, glands separate and oblong, apex entire, rounded. Stamens ca. 4 mm; anther connective slightly prolonged, rounded. Ovules 6, subsessile. Berry shiny, red, subglobose, 10-12 × 9-12 mm, not pruinose, style persistent or not. Fl. May-Jun, fr. Jul-Aug.

Sparse forests, thickets, grasslands; 3500-4500 m. Qinghai, Xizang [NE India, Nepal].